REVSYS: SYSTEMATICS OF THE
SCORPION FAMILY VAEJOVIDAE
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FAMILY VAEJOVIDAEGenus ParavaejovisGenus ParuroctonusGenus PseudouroctonusGenus SerradigitusGenus SmeringerusGenus SyntropisGenus UroctonitesGenus UroctonusGenus Vaejovis

      eusthenura group
           Vaejovis bilineatus
           Vaejovis coahuilae
  
        Vaejovis confusus
   
       Vaejovis diazi diazi
    
      Vaejovis diazi transmontanus
   
       Vaejovis eusthenura
   
       Vaejovis flavus
    
      Vaejovis galbus
    
      Vaejovis glabrimanus
    
      Vaejovis globosus
   
       Vaejovis gravicaudus
  
        Vaejovis hoffmanni hoffmanni
   
       Vaejovis hoffmanni fuscus
    
      Vaejovis punctatus punctatus
   
       Vaejovis punctatus spadix
    
      Vaejovis punctatus variegatus
   
       Vaejovis puritanus
    
      Vaejovis spinigerus
   
       Vaejovis viscainensis
  
        Vaejovis vittatus
  
        Vaejovis waeringi
   
       Vaejovis waueri
      intrepidus group
  
        Vaejovis intrepidus intrepidus
  
        Vaejovis intrepidus atrox
  
        Vaejovis intrepidus cristimanus
    
      Vaejovis occidentalis
    
      Vaejovis subcristatus
      mexicanus group
  
        Vaejovis carolinianus
  
        Vaejovis cashi
  
        Vaejovis chamelaensis
    
      Vaejovis chiapas
   
       Vaejovis chisos
  
        Vaejovis deboersi
  
        Vaejovis dugesi
  
        Vaejovis feti
  
        Vaejovis franckei
   
       Vaejovis granulatus
    
      Vaejovis jonesi
   
       Vaejovis lapidicola
    
      Vaejovis maculosus
   
       Vaejovis mexicanus mexicanus
    
      Vaejovis mexicanus smithi
    
      Vaejovis monticola
    
      Vaejovis nigrofemoratus
  
        Vaejovis pattersoni
   
       Vaejovis paysonensis
    
      Vaejovis pusillus
   
       Vaevovis rosssmanni
    
      Vaejovis setosus
    
      Vaejovis sprousei
    
      Vaejovis tesselatus
    
      Vaejovis vaquero
  
        Vaejovis vorhiesi
      nitidulus group    
    
      Vaejovis curvidigitus
    
      Vaejovis davidi
    
      Vaejovis decipiens
    
      Vaejovis gracilis
    
      Vaejovis intermedius
    
      Vaejovis janssi
    
      Vaejovis kochi
    
      Vaejovis mauryi
    
      Vaejovis minckleyi
    
      Vaejovis mitchelli
    
      Vaejovis nigrescens
    
      Vaejovis nitidulus
    
      Vaejovis norteno
    
      Vaejovis peninsularis
    
      Vaejovis platnicki
    
      Vaejovis pococki
    
      Vaejovis rubrimanus
    
      Vaejovis solegladi
      punctipalpi group
    
      Vaejovis bruneus bruneus
    
      Vaejovis bruneus loretoensis
    
      Vaejovis bruneus villosus
    
      Vaejovis cazieri
    
      Vaejovis crassimanus
    
      Vaejovis hirsuticauda
    
      Vaejovis insularis
    
      Vaejovis magdalensis
    
      Vaejovis punctipalpi punctipalpi
    
      Vaejovis punctipalpi barbatus
    
      Vaejovis punctipalpi cerralvensis
    
      Vaejovis russelli
    
      Vaejovis sonorae
      incertae sedis
    
      Vaejovis acapulco
    
      Vaejovis cisnerosi
           Vaejovis kuarapu
    
      Vaejovis mumai
    
      Vaejovis nayarit
    
      Vaejovis pequeno
    
      Vaejovis spicatus
    
      Vaejovis zihuatanejensis
          "Vaejovis" flavescens

Genus Vejovoidus

Catalog of the VaejovidaeVaejovid  Bibliography

Why Study the Vaejovidae?

Vaejovis glabrimanus Sissom & Hendrixson 2005

Vaejovis glabrimanus Sissom & Hendrixson, 2005: 35-39, figs. 1-12.

TYPE(s):  Vaejovis glabrimanus Sissom & Hendrixson, 2005: - Adult male holotype taken from 4.9 mi N La Ascension, Nuevo Léon, México (24°19' N, 99°54' W) on 18 July 1972 by E.A. Liner, R.M. Johnson, and A.H. Chaney; deposited in the Florida State Collection of Arthropods, Gainesville, Florida.

Original description:
Vaejovis glabrimanus Sissom & Hendrixson, 2005: 35-39, figs. 1-12

"Description.—The following description is based on the holotype male; some meristic and morphometric characters of the paratype male are provided in parentheses.
Coloration
. Base color light yellow-brown, almost immaculate. Tergites with paired vestigial dusky p atches along posterior margin. Metasomal segments with carinae yellow orange. Telson yellowish; aculeus dark reddish-brown. Pedipalps: femur and tibia yellow, lighter than body; chela manus yellow, fingers orange-red with underlying dusky pigment; denticles of dentate margin dark brown. Legs pale yellow with some faint dusky markings; tarsi yellowish. Venter: coxosternal region yellowish; pectines yellowish-white; sternites light yellow.
Prosoma
. Carapace approximately as long as wide. Anterior margin essentially straight, with six conspicuous setae; median notch faint, shallow. Entire carapacial surface densely granular. Mesosoma. Tergites I–IV: Median carina on I–II faint, on II–VI weak, smooth to granular; lateral carinae present posteriorly on II–VI, weak, granular. Tergite VII: Median carina weak, smooth, present on anterior one-third; submedian and lateral carinae strong, granulose. Pectinal tooth count (l/r) 15/? (17/17). Sternites III–VI smooth, moderately setose. Sternite VII acarinate.
Metasoma (Fig. 1). Segment I slightly wider than long, II–III slightly longer than wide, V 2.37 (2.40) times longer than wide. Segments I–IV: Dorsolateral carinae on I–III strong, serrate; on IV strong, crenulate; terminal denticles on I–III distinctly enlarged, spiniform. Lateral supramedian carinae on I–III strong, irregularly serrate; on IV moderate, irregularly serrate; terminal denticles distinctly enlarged, spiniform on I–III. Lateral inframedian carinae on I complete, strong, irregularly serrate; on II present only on posterior third, crenulate to serrate; on III represented by only three or four medium-sized granules; on IV absent. Ventrolateral carinae on I moderate, irregularly crenulate; on II–IV moderate to strong, crenulate. Ventral submedian carinae on I–III weak, smooth; on IV weak, smooth anteriorly, serrate posteriorly. Intercarinal spaces mostly smooth, lustrous. Segment V: Dorsolateral carinae moderate, granular on anterior half of segment. Lateromedian carinae present on anterior third, weak, granular. Ventrolateral and ventromedian carinae moderate, crenulate. Intercarinal spaces mostly smooth, lustrous. Metasomal I–IV carinal setation (l/r): dorsolaterals, 0/0:2/2:2/2:4/4; lateral supramedians, 1/1:3/3:3/3:4/4; lateral inframedians, 2/2:0/0:0/0:0/0; ventrolaterals, 2/3:4/4:4/4:4/4; ventral submedians, 3/3:3/3:4/3:4/5 (segment III with unpaired seta offset medially from carina; segment IV with left seta of pair 2 offset medially from carina). Setation of metasomal segment V: dorsolaterals, 7/7; lateromedians, 4/5; ventrolaterals, 10/10.
Telson (Fig. 1). Dorsal surface flattened, smooth; ventral surface almost smooth, without a subaculear tubercle; aculeus much shorter than vesicle.
Pedipalps. Orthobothriotaxic, Type C (Vachon 1974). Femur (Fig. 2) tetracarinate. Dorsointernal, dorsoexternal, and ventrointernal carinae strong, granulose. Ventroexternal carina moderate, with irregularly spaced, large rounded granules. Internal face with large conical granules; dorsal face shagreened. Ratio of femur length/width, 2.55 (2.52). Patella (Fig. 3) tetracarinate. Dorsointernal carina strong, granulose. Ventrointernal carina moderate, granulose. Dorsoexternal carina weak to moderate, smooth. Ventroexternal carina weak, smooth. Internal face with moderate basal tubercle and longitudinal series of 6–7 large granules. Surfaces smooth. Ratio of patella length/width, 2.47 (2.41). Chela (Figs. 4–7): carinae of palm (Figs. 4–5) obsolete or at most, vestigial and smooth. Dentate margin of fixed finger (Fig. 6) with primary row of denticles divided into six subrows by five larger granules; six inner accessory granules of which all but distal-most paired with larger granule in primary row; trichobothrium it slightly basal to sixth inner accessory granule. Dentate margin of movable finger on right pedipalp chela (Fig. 7) with primary denticle row broken up into six subrows by five enlarged granules; apical row consisting of one small granule; seven inner accessory granules, of which distal granule not paired with larger granule in primary row; basal inner accessory granule distinctly basal to corresponding granule in primary row. Dentate margin of movable finger on left pedipalp chela anomalous, with primary row broken into four subrows by three enlarged granules; distalmost subrows fused; only three inner accessory granules. Slight recess indicated on both fingers such that a distinct gap is visible proximally when the fingers are closed (Fig. 5). Ratio of chela length/width 2.94 (3.06); of movable finger length/chela width 1.60 (1.64); of fixed finger length/carapace length ratio 0.56 (0.53). Legs. Tarsomere I on legs I–II with one retrolateral and two ventrolateral rows of spinules; ventral rows complete, interrupted at irregular intervals by large, dark, stiff setae. Retrolateral row present on distal one-half, interrupted by one or two spines. Tarsomere I spinule rows rudimentary on legs III–IV, but with stiff setae present. Tarsomere II on all legs with single ventromedian row of spinules, procurved basally, terminating distally between with 2–3 pairs of medium-sized spines (Fig. 8).
Hemispermatophore
(Figs. 9–12). Hemispermatophore with broad flange on ectal edge of distal lamina (Figs. 9, 11); capsular region as in Fig. 10; "sperm plug" of capsular region with 14 spines (Fig. 12).
Measurements of Male Holotype (mm):
Total L, 40.85; carapace L, 5.00; mesosoma L, 10.50; metasoma L, 19.10; telson L, 6.25. Metasomal segments: I L/W/D, 2.55/2.70/2.15; II L/W/D, 3.05/2.65/2.15; III L/W/D, 3.25/2.65/2.15; IV L/W/D, 4.10/2.55/2.05; V L/W/D, 6.15/2.60/1.90. Telson: vesicle L/W/D, 4.45/2.45/2.10; aculeus L, 1.80. Pedipalps: femur L/W, 3.70/1.45; patella L/W, 4.20/1.70; chela L/W/D, 7.05/2.40/2.45; fixed finger L, 2.80; movable finger L, 3.85.
Measurements of Male Paratype (mm):
Total L, 42.00 (estimated, due to broken aculeus); carapace L, 5.20; mesosoma L, 11.00; metasoma L, 19.65; telson L, ??. Metasomal segments: I L/W/D, 2.60/3.00/2.30; II L/W/D, 3.10/2.80/2.30; III L/W/D, 3.30/2.70/2.30; IV L/W/D, 4.30/2.60/2.20; V L/W/D, 6.35/2.65/2.05. Telson: vesicle L/W/D, 4.60/2.55/2.10; aculeus L, ?. Pedipalps: femur L/W, 3.65/1.45; patella L/W, 4.10/1.70; chela L/W/D, 7.20/2.35/2.45; fixed finger L, 2.75; movable finger L, 3.85.
Variation
. The paratype male did not differ significantly from the holotype in any of the above-mentioned characters. The setation of the carinae of metasomal segments I–IV (l/r) in the paratype is as follows: dorsolaterals, 2/2:2/2:4/4:5/5; lateral supramedians, 1/2:3/4:4/4:4/4; lateral inframedians, 3/2:1/2:1/1:1/1; ventrolaterals, 3/3:4/4:4/4:4/4; ventral submedians, 3/3:4/3:4/3:5/5 (segments II and III each with unpaired seta slightly offset from the ventral submedian carinae; segment IV with setal pairs 2 and 4 slightly offset medially from the ventral submedian carinae). For metasomal segment V (l/r), the setation of the paratype is: dorsolaterals, 9/10; lateromedians, 4/5; ventrolaterals, 10/10."

Subsequent Accounts

Distribution: NORTH AMERICA. México (Coahuila, Nuevo León).

Published Records: Paratype male taken from 10.8 mi S, 0.1 mi E Arteaga at Los Pinos, Coahuila, México (25°17' N, 100°50' W) on 17 July 1975 by E.A. Liner et al.; also deposited in the Florida State Collection of Arthropods.

Notes: In possessing a setose metasoma and essentially acarinate pedipalp chelae, Vaejovis glabrimanus sp. nov. is most similar to V. coahuilae and V. globosus. It may be distinguished from V. coahuilae by (1) the presence of six, rather than five, subrows of denticles on the cutting margin of the pedipalp chela fixed finger; (2) the absence of an elaborate dusky pattern on the carapace, dorsum, metasoma, and appendages; (3) the lack of carinae on sternite VII (in V. coahuilae, granular lateral carinae are present); and (4) larger body size, with adult males reaching approximately 40–42 mm in length (adult males of V. coahuilae are rarely about 35 mm in length). It may be distinguished from V. globosus by (1) the presence of six, rather than five, subrows of denticles on the pedipalp chela fixed finger; (2) the presence of prominent scalloping in the chela fingers in the male; (3) the absence of dusky markings on the dorsum and metasoma; (4) more swollen pedipalp chelae in the male (chela length/width ratios: V. glabrimanus, 2.94–3.06; V. globosus, 3.29–3.79); and (5) its larger body size (males of V. globosus range only to about 25–30 mm). In body size and coloration, the new species is also superficially similar to V. russelli Williams. As is typical of the punctipalpi group, males of V. russelli have distinctly carinate, often granular pedipalp chelae and fewer metasomal setae (e.g., segment V in V. russelli typically has only 4–7 setae on the metasomal dorsolateral and ventrolateral carinae, whereas V. globosus has 7–10 on the dorsolaterals and 10 on the ventrolaterals). Additional characters for the punctipalpi group are provided by Williams (1971b, 1980).The specific epithet is Latin for "smooth hand" and refers to the lack of distinct carinae on the pedipalp chelae.

Sissom and Hendrickson 2005:39,40,41 -  Key to the Vaejovid Scorpion Species of Northeastern México  - Dorsolateral carinae of metasoma (at least on segments I–III) with an enlarged terminal denticle; legs I–III with irregular setation (except in V. globosus, which has setal combs, – but has the enlarged terminal denticle on the metasoma); Trichobothria ib and it situated more distally, at the level of the sixth inner accessory denticle or beyond; Ventral submedian carinae of metasoma developed and usually granular, at least on segments III and IV; Metasoma setose (e.g., metasoma V ventrolateral carinae with 9–12 setae); pedipalp chelae of adult male acarinate or at most feebly carinate; Cutting margin of pedipalp chela fixed finger with 6 subrows of denticles; underside of metasoma uniformly yellowish ..................................Vaejovis glabrimanus sp. nov.

 


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