Pseudouroctonus m. castaneus
Pseudouroctonus m. thompsoni
Pseudouroctonus williamsi (Gertsch & Soleglad 1972)
williamsi Gertsch & Soleglad, 1972: 566, 579-580, fig. 11, 20, 34,
35, 125, 126; Soleglad, 1973b: 353.
Vejovis williamsi: Stahnke, 1974: 130, 136; Williams & Savary, 1991:
Vaejovis angelenus (part): Williams, 1976: 2 (see note below).
Pseudouroctonus williamsi: Stockwell, 1992: 409; Kovarík, 1998: 145;
Gertsch and Soleglad 1972 - Female holotype from Mission Gorge, 1 mi. W
Padre Dam, 100 ft., San Diego County, California, July 7, 1969 (S. C.
Williams, V. F. Lee), in collection of California Academy of Sciences (CAS,
Type No. 11475)..
Gertsch and Soleglad 1972:
"DIAGNOSIS: Large, dusky brown,
heavily granulated species similar to mordax except as follows: average
size about 45 mm. with largest female 54 mm. long; median eye tubercle
lying in front of middle of carapace; pectinal tooth count 11 for
females, 13 for males; lower margin of movable finger of chelicera
smooth; inferior lateral and median keels of basal segments of cauda
prominent and granulated; keels on chela crenulate to heavily
granulated; vesicle of sting considerably inflated at apex.
COLORATION: Of fully adult specimens as follows: carapace dark reddish
brown, with granules on keels and eye tubercles blackish; preabdomen and
cauda dusky brown; pedipalps dark reddish brown, quite shiny, with keels
and fingers blackish; legs, sternites of preabdomen, chelicerae, and
telson, except dark sting, yellowish. Immature specimens yellowish, with
reddish brown pedipalps.
STRUCTURE: Similar in both sexes to that of mordax in basic features
unless otherwise indicated. For measurements, see table 4.
Carapace: Entire surface covered thickly with large, round granules;
coriaceous, with median grooves and posterior side depressions prominent
and bordered by submarginal trench; frontal lobes with six stiff
bristles. Frontal emargination a deep, rounded notch. Median eyes
situated back about three-eighths of distance from front to posterior
margin. Median eyes small, 0.34 mm., separated by full diameter. Lateral
eyes (fig. 20) three, two large ones close together overhanging side
margin; third eye small, inconspicuous, placed above second eye.
Preabdomen: All tergites quite coarsely granulated, except along basal
strip, with heaviest granules along posterior edges; distinct traces of
median keels present as rows of heavier granules on midline of tergites
I-VI. Tergite VII granulose, with dorsal and lateral keels prominent,
each set with heavy, rounded granules, and with many large granules in
intercarinal spaces. Sternites smooth, shiny; sternite V with sides
granulose and development of weak carinae in males, these much reduced
in females. Shape of stigma ofmale as shown in figure 11.
Cauda: In both sexes widest at base and slightly narrowed at segment V.
All caudal segments longer than wide; segment V, slightly longer than
carapace. Keels on all surfaces prominent and granulated to give
serrated appearance in lateral view; dorsal and superior lateral keels
showing no prominent development of apical granules at posterior ends of
series. Bristles on cauda short; three principal ventral pairs on all
Telson: See figures 125, 126. Vesicle suboval, broader than fifth caudal
segment, flattened above and inflated behind near sting, smooth above
but with fine granulation below and on sides, provided with a few short
bristles. Sting one-third as long as vesicle.
Pectines: Those of female: median piece twice as broad as long, with
shallow, V-shaped indentation on frontal margin; pectin length less than
twice that of median piece (40/23); middle lamellae, seven or eight;
pectinal teeth, 11. Those of male with smaller, deeply indented median
piece, which is about twic San
e as long as pectin (70/38); middle
lamellae, six; pectinal teeth, 13.
Genital opercula: Like those of mordax.
Chelicerae: See figures 34, 35. Basic pattern like that of mordax but
teeth short, rounded; lower margin ofmovable finger a smooth keel.
Pedipalps: In both sexes like those of mordax except as follows: frontal
spurs on tibia of medium size, with prominent, pointed granules on each
side bearing short seta; all keels on chela prominent, all granulated;
superior keel finely crenulated as seen in lateral view; intercarinal
spaces finely granulated above, smoother below.
Walking legs: Setation like that of mordax; tarsus with three thin
spines on each side of ventral line of spinules."
AMERICA. USA: (California:San Diego County).
California: San Diego County:
Mission Gorge, 1 mi. W Padre Dam, 100 ft., July 7, 1969 (S. C. Williams, V.
F. Lee), three males and three immature females, in collection of California
Academy of Sciences; Mission Gorge, March 31, 1940 (Earl Brown), female, in
San Diego Museum; Wild Cat Canyon, August 9, 1962 (S. C. Williams), female
probably this species from pit trap in burned area.
Gertsch and Soleglad (1972)
dedicated this species "to Dr. Stanley C. Williams, San Francisco State
College, who has made numerous contributions to knowledge ofAmerican
scorpions. Williams (1976) listed V. williamsi (Gertsch &
Soleglad, 1972) as a junior synonym of V. angelenus (Gertsch &
Soleglad, 1972). Later, Williams & Savary (1991) regarded them as separate