REVSYS: SYSTEMATICS OF THE
SCORPION FAMILY VAEJOVIDAE
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FAMILY VAEJOVIDAEGenus ParavaejovisGenus ParuroctonusGenus Pseudouroctonus
     Pseudouroctonus andreas
     Pseudouroctonus angelenus
     Pseudouroctonus apacheanus
     Pseudouroctonus bogerti
     Pseudouroctonus cazieri
     Pseudouroctonus chicano
     Pseudouroctonus glimmei
     Pseudouroctonus iviei
     Pseudouroctonus lindsayi
     Pseudouroctonus minimus
          Pseudouroctonus m. minimus
          Pseudouroctonus m. castaneus
          Pseudouroctonus m. thompsoni
     Pseudouroctonus reddelli
     Pseudouroctonus rufulus
     Pseudouroctonus
savassi
     Pseudouroctonus sprousei
     Pseudouroctonus williamsi

Genus SerradigitusGenus SmeringerusGenus SyntropisGenus UroctonitesGenus UroctonusGenus VaejovisGenus Vejovoidus

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Pseudouroctonus reddelli (Gertsch & Soleglad 1972)

Vejovis reddelli Gertsch & Soleglad, 1972: 595-598, fig. 12, 22, 38, 39, 135; Soleglad, 1973b: 359.
Vejovis
sp.: Reddell, 1965: 167; Reddell, 1970: 402.
Pseudouroctonus reddelli
: Stahnke, 1974: 132; Kovarík, 1998: 145; Sissom, 2000:517-518; Soleglad & Fet, 2003b: 6, fig. 7; Soleglad & Fet, 2003a: 8, 28, 33, 58, 67, 103, 162, figs. 6, 36, 61, 73; Soleglad & Fet, 2005: 2, 6; Prendini & Wheeler, 2005: 479.
Vaejovis reddelli
: Reddell, 1981: 114; Sissom, 1990a: 109; Sissom & Jackman, 1997: 151, 155.

type(s):  Vejovis reddelli Gertsch & Soleglad 1972 - Female holotype from Pablo's Cave, Uvalde County, Texas, April 5, 1963 (J. Reddell, D. McKenzie), found in first room of cave in darkness about 50 feet from entrance, deposited in the American Museum of Natural History.

Original Description:
Gertsch & Soleglad, 1972:

"DIAGNOSIS: Large, dark scorpion resembling mexicanus and related species of Mexico, readily distinguished by following features: all segments of cauda with ventral keels granulated; first caudal segment slightly broader than long in females, as long as broad in males, succeeding segments usually longer than broad in both sexes; keels on chela rather weakly developed and granulated; keel on lower margin of movable finger of chelicera with several dark teeth; pectinal teeth of females 15-16, of males 18-19.
...
COLORATION: General appearance in both sexes that of large, reddish to mahogany brown, or blackish scorpion, with tan walking legs and chelicerae. Underside mostly tan, with pectines yellowish. Young specimens paler, with dusky pattern.
STRUCTURE: Dorsal view of male as shown in figure 135. Similar in both sexes, except for smaller size of males. Measurements as given in table 9.
Carapace: Coriaceous, with medium-sized granules in females, concentrated in posterior half and larger ones in males. Area around median eye tubercle and interocular triangle quite smooth in females, rugulose and with heavier granules in males. Frontal margin with rather deep-rounded indentation and set with six long setae; this margin in males revolved to form rugose ridge armed with row of granules. Median eyes about 0.32 mm., separated by more than their diameter, on rather prominent tubercle; eye tubercle about one-fifth width of carapace at that point and situated in front of middle point of length. Lateral eyes (fig. 22) three; posterior eye only slightly smallest.
Preabdomen: Tergites I-II essentially smooth at base, with small granules along posterior edges; tergites III-VI smooth at base but with posterior half or more of each with scattered granules; tergite VII with scattered granules and prominent serrate keels. Stigma of sternite IV of male as shown in figure 12.
Cauda: Dorsal and superior lateral keels sharply serrate, each terminating in sharp, erect spine. Lateral keels serrate on caudal segment I, present and serrate on posterior quarter of segment II, serrate on posterior fifth ofsegment III, and absent on segment IV. Inferior lateral keels serrate on segments I-IV. Inferior median keels unevenly smooth on segment I, weakly serrate on segment II, serrate on segments III-IV. Intercarinal spaces essentially devoid of granulations. Dorsal keels on segment V with uneven patches of weakly serrate granules; lateral keels weak on anterior half, formed by small, pointed granules; inferior lateral keels strongly serrate; single median keel with serrate granules. Ventral setae count of segments I-V, 3-3-3-3-4.
Telson: Vesicle wider than fifth caudal segment in both sexes. Sting about half as long as vesicle, rather abruptly curved at apex, with
inconspicuous nodule and two projecting setae at base.
Pectines:
Median piece about two-thirds as long as wide, with shallow emargination in front.
Median lamellae circular, eight to 12 in both sexes. Pectinal teeth short in females, longer and angular in males; counts of females 15-16, of males 18-19.
Chelicerae: See figures 38, 39. Fixed finger of standard vejovid form with respect to count and position; two or three teeth or nodules present on inconspicuous ventral carina. Movable finger with normal five strong teeth on upper margin; lower margin with heavy distal tooth and keel bearing four or five strong, sharp or rounded, dark teeth. Serrulae present on lower edge of movable finger.
Pedipalps: Similar in both sexes with heavy hands and long fingers. Femur with all keels sharply serrate. Tibia with inner keels serrate, outer keels crenulate, and raised portion on inner face set with two blunt spines. Chela with all keels weakly defined and with no sharply marked ridges. Granulation of superior keel quite smooth, with uneven, rounded granules; inner accessory keel smooth; outer accessory keel barely distinguishable, covered with minute granules; inner keel rounded, with heavy, uneven granules. Inferior keel rounded, with medium-sized granules; inner ventral keel obsolete; inner accessory keel weakly granulated. Six files of fine teeth on inner carinae of fingers, flanked by six supernumerary teeth on fixed and seven on movable finger.
Walking legs: Tarsi with median line of ventral spinules flanked by four or five setae on each side."

subsequent accounts: Sissom and Hendrickson 2005:39,40 Key to the Vaejovid Scorpion Species of Northeastern México  - Dorsolateral carinae of metasoma (at least on segments I–III) with an enlarged terminal denticle; legs I–III with irregular setation (except in V. globosus, which has setal combs, – but has the enlarged terminal denticle on the metasoma); Trichobothria ib and it situated at base of chela fixed finger; Pectinal tooth counts greater than 14 in males, greater than 11 in females; pedipalp chela with ventral face rounded, with or without ventromedian carina; chela movable finger length/ chela palm width ratio greater than 1.8; adults variable in size and color; Cheliceral fixed finger with a carina bearing denticles along the ventral margin; Pectinal tooth counts less than 20 in both sexes; cheliceral movable finger with distinct denticles along ventral margin; adults dark reddish brown in color ................ Pseudouroctonus reddelli

distribution: NORTH AMERICA. USA (Texas). Per Gertsch and Soleglad (1972) , Reddell (1965, p. 167) listed 22 caves in 10 counties where specimens have been taken and classified the species as a "common troglophile in caves throughout the state." Specimens from Texas caves previously referred to Vejovis mexicanus and crassimanus probably are this species. The species also occurs outside of cave habitats as shown by some of the following records.

Published Records:  Texas: Bandera County: Fog Fissure, 5 mi. N Vanderpool, October 30, 1963 (D. McKenzie), female. Bexar County: 1 mi. N Helotes, July 2, 1959 (J. F. Lawrence), male; Government Canyon Bat Cave, 5 mi. SW Helotes, August 11, 1965 (J. Reddell, J. Fish), male; Madla's Ranch Cave, 3 mi. N Helotes, July 6, 1959 (J. F. Lawrence, J. Reid), male; Adam Wilson's Cave, 4 mi. N Helotes, October 4, 1963 (J. Reddell, D. McKenzie), juvenile. Burnet County: Pie Cave, X mi. N Longhorn Caverns, July 24, 1963 (W. Russell), immature. Comal County: New Braunfels, May 28, 1918, female; Brehmer Cave, New Braunfels, September 11, 1940 (W. Jellison, G. Kohls), six males, six females. Edwards County: Deep Cave, W mi. E Punkin Cave, September 4, 1965 (J. Reddell, D. McKenzie), male, two females from walls of upper room; September 4, 1965 (D. Dickey), two immatures; Jacoby Cave, about 10 mi. W Telegraph, just south of Kimble County line, September 22, 1963 (J. Reddell, D. McKenzie), young female. Hays County: Ezell's Cave, San Marcos, September 7, 1963 (J. Reddell, D. McKenzie, R. Ballinger), male; Donaldson Cave, August 15, 1965 (B. Frank, B. Banfer), male. Kendall County: Near Century Caverns, 11 mi. NE Boerne, March 20, 1964 (W. Russell), female from under rock near cave entrance. Kerr County: Between Hunt and Leakey, December 30, 1956 (W. McAlister, K. Baker), young female; 6 mi. NW Hunt, December 29, 1956 (W. McAlister, K. Baker), female. Kimble County: Fleming Bat Cave, 2 mi. N Telegraph, September 3, 1967 (J. Goodbar), female, with 21 young. Real County: Orell Crevice Cave, 100 yards W Orell Bat Cave, August 18, 1963 (J. Reddell, D. McKenzie), immature on wall; Skeleton Cave, 5 mi. N Leakey, August 18, 1963 (J. Reddell, D. McKenzie), male, young female from under rocks in entrance room. Travis County: Beckett's Cave, 1 mi. S Oak Hill, December 5, 1965 (W. Russell), female; Cotterrell Cave, 7 mi. N Austin, March 10, 1967 (E. E. Remington), two males; Bee Creek Cave, west of Tom Miller Dam in Austin, October 2, 1963 (J. Reddell, D. McKenzie), young female; Ken Harrell Cave, 10 mi. N Austin, March 27, 1963 (W. Russell), three immatures; Broken Straw Cave, Austin, March 10, 1967 (E. E. Remington), male; Bandit Cave, Austin, March 8, 1968 (J. Reddell, J. Fish), male; Goat Cave, 8 mi. SW Austin, March, 1968 (J. Reddell, J. Fish); Kretschmarr Cave, 15 mi. NW Austin, March 2, 1963 (J. Reddell, D. McKenzie), one immature from second room, 30 ft. from entrance; Cave X, 6 mi. S Austin (J. Reddell), young male, March 8, 1963 (J. Reddell, D. McKenzie), female; Tooth Cave, 15 mi. NW Austin, October, 1963 (W. Russell), female, February 25, 1963 (D. McKenzie, W. Russell), male, August 5, 1963 (J. Reddell), immature from under rocks; 2 mi. W Austin, April 2, 1958 (W. McAlister), female; Bull Creek, 8 mi. NW Austin, August 22, 1968 (B. Vogel). Zilker Park, Austin, April 1, 1956 (W. McAlister, J. Boyd), male. Uvalde County:  North Well Cave, April 5, 1963 (J. Reddell, D. McKenzie), two young males from anteroom and main cave room; McNair Cave, October 26, 1965 (J. Reddell, J. Fish, J. Calvert), female; Tampke Ranch Cave, February 11, 1966 (J. Reddell, D. McKenzie), male. Val Verde County: 6 mi. N Del Rio, April 10, 1968 (J. Reddell), two immatures. Williamson County: Williams Cave, 2 mi. NW Georgetown, August 24, 1968 (J. Reddell, W. Russell), immature; Steam Cave, 2 mi. S Georgetown, July 7, 1963 (J. Reddell, W. Russell), male from wall in darkness; Cave 31, October, 1964, male; Inner Space Caverns, 2 mi. S Georgetown, December 22, 1968 (W. Elliott), male; Core Hole Cave, 1 mi. S Georgetown, November 3, 1963 (J. Reddell, D. McKenzie, J. Porter), two males; Steam Cave, 2 mi. S Georgetown, January 6, 1963 (J. Reddell, T. Phillips), three immatures.

notes: Gertsch and Soleglad (1972) named this species "for Mr. James Reddell, student of caves and cave faunas, who has done extensive exploration of Texas and Mexican caves.".

 


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